Agnostus
| Agnostus |
|
|---|---|
| File:Agnostus pisiformis Kinnekulle 1 CRF.jpg | |
| disarticulate cephalons and pygidia of Agnostus pisiformis, from Kinnekulle, Sweden | |
Scientific classification  |
|
| Kingdom: | Animalia |
| Phylum: | Arthropoda |
| Class: | †Trilobita |
| Order: | †Agnostida |
| Family: | †Agnostidae |
| Genus: | †Agnostus Brongniart, 1822 |
| subgenera | |
and see text |
|
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Agnostus is a genus of agnostid trilobites that lived during the upper Middle Cambrian–lower Upper Cambrian (approximately 506 to 492 million years ago). It is the type genus of the family Agnostidae. It is subdivided into two subgenera, Agnostus and Homagnostus.[1][2]
Contents
Etymology
Agnostus is from the Ancient Greek άγνωστος (ágnostos) "unknown", and reflects the fact that early paleontologists did not know what type of animal they were confronted with. The species epithets have the following origins:
- pisiformis derives from the Latin pīsum which means "pea", and forma meaning "shape", for its globe-like appearance when rolled.
Description
Like all Agnostida, Agnostus is diminutive and the headshield (or cephalon) and tailshield (or pygidium) are of approximately the same size (or isopygous) and outline. As in all Agnostina, Agnostus has two thorax segments. When Agnostina are enrolled, an tiny opening between the cephalon and the thorax becomes visible (the cephalothoracic aperture) that uniquely distinguishes this group from the Eodiscina. Two features shared with all Agnostoidea are that the basal glabellar lobes are anteriorly expanded, and that the frontal lobe of the central raised area of the cephalon (or glabella) is not laterally expanded compared to the backward lobe. This is unlike in the Condylopygoidea. Like in all Agnostidae, the cephalic border is narrow. Like in all Agnostinae, the pygidial axis is simple, the posterior lobe not wider than the anterior (or axiolobate), and usually long but not reaching border furrow. In Agnostus the glabella is connected to the border by a median preglabellar furrow. The anterior glabellar lobe is ogival or rounded at its front. The posterior lobe is tapering forward. The glabella carries an axial node level with or slightly behind a variably developed second furrow. The pygidium has a long axis, although this usually does not reach the border furrow. The pygidial axis (or rhachis) is not connected to the border by a median (postaxial ) furrow. The pygidium carries backward directed spines on its margins, where it curves back towards the midline.
In the subgenus Agnostus (Agnostus), the axis of the pygidium is relatively narrow, ending pointed, or narrowly rounded, and furrows crossing the pygidial axis are weak at best. The subgenus Agnostus (Homagnostus) has a broadly rounded termination of the axis extending nearly to posterior border furrow. The axis is narrower halfway along its length and crossing furrows are well developed.[4]
Appendages
The appendages of meraspid developmental stages of Agnostus pisiformis (½-1 mm total body length) are the best preserved of any trilobite found sofar.[4] Agnostus pisiformis has nine pairs of appendages, the antennules, and eight pairs of "legs", that differ in size and composition. Four pairs of appendages originate from the cephalon, two from the thorax and three from the pygidium. The frontal antennules, are short and stout. The most forward pair of "legs" has a small coxa, a 7 podomere exopodite carrying long bristle-like setae and gill-like extensions, and no endopodite. The second pair has a 5 podomere exopodite and a 6 podomere endopodite. The third a 3 podomere exopodite and a 6 podomere endopodite. The fourth a 2 podomere exopodite and a 7 podomere endopodite. The fifth a single podomere exopodite and a 6 podomere endopodite. Further appendages are more and more reduced in size and complexity.[5]
Distribution
Agnostus is said to have had a cosmopolitan distribution. The nominate subgenus occurs from the late Middle Cambrian to the early Upper Cambrian, while the subgenus Agnostus (Homagnostus) is restricted to the early Upper Cambrian.
- A. artilimbatus occurs in the Middle Cambrian of Kazakhstan (Kyrshabakty River: Lejopyge armata trilobite zone, Zhumabai Formation, Goniagnostus nathorsti trilobite zone, Aktas Formation; Ayusokkanian (505.0 - 498.5 Ma), 43.5° N, 70.0° E).[6][7]
- A. pisiformis is known from the earliest Upper Cambrian of Sweden (A. pisiformis-zone, Alum shale, Adrarum, Scania).
Taxonomy
Agnostus pisiformis was the first nominal species to be described and illustrated in the literature. Magnus von Bromell in 1729 described disarticulated heads and tails in matrix as "minimorum vermiculorum vaginipennium" ("small beetle-like worms").[8] The earlier concepts of the genus Agnostus were very wide, initially including almost all of the Agnostida, but this has gradually been narrowed down. This is the reason why a large number of species are now assigned to other genera, often in other families or even to the Eodiscina.
Species
Species classified under the genus include the following:[1][2]
Column-generating template families
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| Type | Family |
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Responsive/ Mobile suited |
Start template | Column divider | End template |
|---|---|---|---|---|---|---|
| Float | "Col-float" | Yes | Yes | {{Col-float}} | {{Col-float-break}} | {{Col-float-end}} |
| "Columns-start" | Yes | Yes | {{Columns-start}} | {{Column}} | {{Columns-end}} | |
| Columns | "Div col" | Yes | Yes | {{Div col}} | – | {{Div col end}} |
| "Columns-list" | No | Yes | {{Columns-list}} (wraps div col) | – | – | |
| Flexbox | "Flex columns" | No | Yes | {{Flex columns}} | – | – |
| Table | "Col" | Yes | No | {{Col-begin}}, {{Col-begin-fixed}} or {{Col-begin-small}} |
{{Col-break}} or {{Col-2}} .. {{Col-5}} |
{{Col-end}} |
Can template handle the basic wiki markup {| | || |- |} used to create tables? If not, special templates that produce these elements (such as {{(!}}, {{!}}, {{!!}}, {{!-}}, {{!)}})—or HTML tags (<table>...</table>, <tr>...</tr>, etc.)—need to be used instead.
Species previously assigned to Agnostus
As the genus Agnostus was erected early on, many species have since been reassigned.[4][9]
Column-generating template families
The templates listed here are not interchangeable. For example, using {{col-float}} with {{col-end}} instead of {{col-float-end}} would leave a HTML "div" (division) open, potentially harming any subsequent formatting.
| Type | Family |
Handles wiki
table code? |
Responsive/ Mobile suited |
Start template | Column divider | End template |
|---|---|---|---|---|---|---|
| Float | "Col-float" | Yes | Yes | {{Col-float}} | {{Col-float-break}} | {{Col-float-end}} |
| "Columns-start" | Yes | Yes | {{Columns-start}} | {{Column}} | {{Columns-end}} | |
| Columns | "Div col" | Yes | Yes | {{Div col}} | – | {{Div col end}} |
| "Columns-list" | No | Yes | {{Columns-list}} (wraps div col) | – | – | |
| Flexbox | "Flex columns" | No | Yes | {{Flex columns}} | – | – |
| Table | "Col" | Yes | No | {{Col-begin}}, {{Col-begin-fixed}} or {{Col-begin-small}} |
{{Col-break}} or {{Col-2}} .. {{Col-5}} |
{{Col-end}} |
Can template handle the basic wiki markup {| | || |- |} used to create tables? If not, special templates that produce these elements (such as {{(!}}, {{!}}, {{!!}}, {{!-}}, {{!)}})—or HTML tags (<table>...</table>, <tr>...</tr>, etc.)—need to be used instead.
References
- ↑ 1.0 1.1 Lua error in package.lua at line 80: module 'strict' not found.
- ↑ 2.0 2.1 Morphology, Ontogeny, and Life Habit of Agnostus pisiformis From the Upper Cambrian of Sweden - Number 19
- ↑ Lua error in package.lua at line 80: module 'strict' not found.
- ↑ 4.0 4.1 4.2 Whittington, H. B. et al. Part O, Treatise on Invertebrate Paleontology. Revised, Volume 1 – Trilobita – Introduction, Order Agnostida, Order Redlichiida. 1997
- ↑ Lua error in package.lua at line 80: module 'strict' not found.
- ↑ Lua error in package.lua at line 80: module 'strict' not found. cited in Lua error in package.lua at line 80: module 'strict' not found.
- ↑ Lua error in package.lua at line 80: module 'strict' not found.
- ↑ Lua error in package.lua at line 80: module 'strict' not found.
- ↑ Lua error in package.lua at line 80: module 'strict' not found.
External links
Data related to Agnostus at Wikispecies
Media related to Lua error in package.lua at line 80: module 'strict' not found. at Wikimedia Commons
